HTR AC VOLTAGES on OUTPUT TUBES are pin 4 (Blk)to pin 5 (RED). The Only DC HTR"S are V-7,V-8, V-9, V-10. These get checked with RED on pin 4 or 5, and Blk on Chassis. They should be in spec when running. Set your variac so you are getting 117AC on the terminal strip side of the CL-80. Then check the output tube AC HTR's (should be 6.3 +/-5%) and V7 to V10. These will have 12VDC on them. You;'l get one of the two scenario's as the tubes are in Series parallel. One tube on each pair will show 25.2V on one pin and 12.6V on the other Pin (for a 12.6V drop) The other tube will show 12.6vdc on one pin and 0.0vdc on the other pin. Allow +/-5%.
Vary the Wall/Variac voltage until both AC and DC Htr voltages are in spec. Note the Voltage. If you don't mind a varying voltage (which the bucking transformer will do based on wall voltage) then a bucking transformer will vary input voltage based on what the Wall voltage is at the moment. And it's fine. If you have gas with varying voltages keep the Variac set for the voltage needed to maintain the heaters at spec. Plate and Screen voltages can vary as much as 50-60 volts either way depending on Wall Voltage, Dynamics vs. Volume, etc. You plate voltages are fine considering the wall voltage on an early 400. Remember that the 400 was designed to operate on Voltages of 105VAC to 120VAC. About a 15% differential. Fisher spec'ed a 15% differential in Voltages on the schematics based on component tolerances. The voltages on the schematic are a guide, and are ideal test voltages based on a highly regulated 117VDC. I run my 400's, 800c's and consoles on CL-80's and wall voltges btwn 120 and 124. No problems over the last 10 years.
Your voltages "IDEALLY" @ 117V input should be PLATE 395VDC, Screen 345VDC, grid -15Vdc(based on OLD STOCK Tubes). Of the nearly a dozen 400's I've rebuilt, NONE of them have Plate, Screen or Grid voltages that matched the Schematic @ 117VAC Wall Votlage. Most have been less than 5% high or low. Close is Good enough in These, Hand grenades and Golf.