My guess, for what its worth....
Terry, good job of eliminating what's *not* wrong.
I'd also bet that there are resistors out of spec in that amp. The last ones I recently re-did had well over 50% of the resistor compliment out of spec with more than one with hairline cracks in the casing.
Also, I'd not overlook the tube socket connectors as a source of noise. If they're loose, they can cause all sorts of gremlins, too. At the age of these amps, they really need to be looked at to remain stable and safe for operation, IMHO. Once done, though, they'll sing like they did as they left the factory back when Kennedy was in office.
Cheers,
David