Even so, the 12au7 plates are still way too high. I'll re-measure and post, but something like a90 on pin one and 260 on pin 6.
Do you mean 190V on pin 1?
I ran a load line for the fist stage of this amp as per the voltage readings and resistor sizes posted on the schematic. I was thinking it might be a misprint, but no, the schematic is smack dab on the money for the first stage.
So there are only a few things that can be wrong:
1. rail voltage not as expected
2. plate resistor not as expected
3. cathode resistor not as expected
4. filament voltage not as expected
5. DC on the grid where none is expected
6. bad tube
I think so far we've ruled out 1 and 4. Or have we ruled out #1? What is the exact rail voltage (voltage at the top side of the 100K resistor with respect to ground) while the amp is idling? Otherwise, that leaves 2, 3, 5, and 6 as possible suspects.
You might want to temporarily ground the grid (pin #2) of the first 12AU7 triode with an alligator clip straight to the chassis so you can guarantee it's at ground level while you measure the other items. Specifically measuring rail voltage, voltage drop across the 100K plate resistor, and voltage drop across the 820 ohm cathode resistor will give you what you need. There is probably something amiss in one of those three. Could be a bad tube. Ruling out all other possibilities would leave a bad tube as the only other option.