I don't much care for those little round transistors, some call them flying saucer types, they sometimes fail and get noisy.
Gain matching may or may not be required in this case.
According to the relevant data sheets;
2SA493 Y gain code is 120 to 240 Hfe.
2SA493 GR gain code is 200 to 400 Hfe.
2SC1000 GR gain code is 280 Hfe
2SC1000 BL gain code is 530 Hfe
If I haven't made any errors, generally speaking, on this style preamp-tone board;
H401,402, 405, 406, (NPN) can be replaced with 2SC1000 (GR or BL) or KSC1845 F gain code.
H403,404, 407, 408, (PNP) can be replaced with 2SA493 (Y or GR) or KSA992F F gain code.
Japanese transistors are usually coded as 2SA, 2SB are PNP, 2SC, 2SD are NPN
Fairchild (and now On-Semi) carried this forward in a way, KSA, KSB are PNP, KSC, KSD are NPN.
YMMV
Tom