If the current readings taken at the cathode test points cannot be brought up to the proper level, then there are four basic circuit areas where the problem could lie -- understanding that the problem could be in any one area, or a combination of them:
1. Low Screen Grid Voltage (pins 4&8 on each output tube). The screen voltage sets the current flow capability of the output tubes at any given control grid (pin 6) bias voltage level. If you are measuring an honest ~ 400 vdc between the screen grid terminals and ground, then this is not the problem.
2. The value of the 10 ohm cathode resistors. If these are honestly 10.0 ohms as measured from pin 5 (the cathode) to ground, then a .22 volt reading across them in fact indicates a current flow of only 22 ma. However, if the 10 ohm resistors should really read only 5 ohms (for example), then current flow across the resistors is actually 44 ma, while you are still "thinking" it's only 22 ma. The value of these resistors must be verified at pin 5 of each output tube.
3. A problem in the bias circuit. Making the voltage more negative at pin 6 of the output tubes only causes the output tubes to draw less current. Jumping the 68K resistors did just that, which should have made the problem worse, and it did. For this exercise, the output tubes could care less HOW the bias voltage is developed and delivered. They just know they want to see about -16 vdc (or so) at pin #6 to produce the target ball park current draw based on the other voltages present. Whatever accomplishes producing (about) this amount of negative voltage at pin 6 of each output tube should cause them to draw the correct amount of current. Since there is (potentially) too much bias voltage, I would be more inclined to double check the value of the 33K resistors at the bias pots, and the 100K resistors at the balance pots. If any of these are significantly high in value, then that would tend to produce the problem you are having.
4. Low heater voltage. If the AC voltage across pins 2&7 are lower than 6.0 vac, then current flow can be a problem if weak tubes are installed. Check the voltage across these terminals with the amplifier biased as close to normal operation as possible.
If ALL of these areas are verified to be as they are supposed to be, then the only other possibility left is that of the tubes themselves. However, since the problem seems to be present equally in both channels, it implies a mistake common to both channels, or a similar mistake made in both channels.
I have no doubt that you will find the issue somewhere within these possibilities. Good luck with it!
Dave