There's a white bottle of "something" near the cab of the truck. That bottle has a hose that looks to be hanging to the floor of the bed. Driving, shaking, bumps and heat caused whatever liquid is in that bottle of "something" to siphon and seep onto the moving blanket beneath the speakers and into the groove of the bed liner where it wicked down to the rear of the bed saturated the moving blanket and was ignited by ?? Possibly mixing with the contents of the other bottle of "stuff" being crushed by the tie down strap. The point of ignition and the hot spot appears to be closest to the tailgate and between the speakers where flames began to follow the chemical (fuel) trail up the moving blanket to its source... the white bottle with "something" in it.
I'd bet that whatever is in the white bottle is "something" flammable. The moving blanket and speakers didn't just catch on fire from a cigarette alone, there is obviously a fuel source involved. IMHO