Huzah!
It lives!
I have a very stout 10 WPC tube amp, that's ruler flat from at least 20hz-20kHZ (didn't measure above or below that) and the voltage is right on the target of 380 at the first step, 335 at the second, and I forget what the third step was at, but it's all within about ±5V which is pretty good.
I had to put in a different rectifier tube, and it works with a bucking transformer to get the voltage just right, and the rectifier takes a while to warm up so by the time it has turned on, so have the rest of the tubes and the power supply stays below 450V the whole time.
Anyway, I just need to make everything that was clipped in externally internal, and then button it up with the final set of tubes and should be good to go!